/* Aşama 2 — yalnızca Biyoloji sayfalarına ait stiller. Ana sayfa stillerini değiştirmez. */
.page-breadcrumb { position: relative; z-index: 2; padding-top: 20px; }
.page-breadcrumb ol { display: flex; flex-wrap: wrap; align-items: center; gap: 9px; margin: 0; padding: 0; list-style: none; color: #73736f; font-size: 10px; }
.page-breadcrumb li + li::before { margin-right: 9px; color: #4d4d49; content: "/"; }
.page-breadcrumb a { transition: color .2s ease; }
.page-breadcrumb a:hover { color: #f4f4f2; }
.page-breadcrumb [aria-current="page"] { color: #c5c5c1; }

.bio-hero { position: relative; overflow: hidden; padding: 46px 0 42px; }
.bio-hero::before { position: absolute; top: -240px; right: -120px; width: 650px; height: 650px; border: 1px solid rgba(255,255,255,.027); border-radius: 50%; box-shadow: 0 0 0 68px rgba(255,255,255,.011), 0 0 0 136px rgba(255,255,255,.006); content: ""; pointer-events: none; }
.bio-hero__grid { position: relative; display: grid; grid-template-columns: minmax(0,1.04fr) minmax(350px,.96fr); align-items: center; gap: clamp(36px,6vw,78px); }
.bio-hero__copy { position: relative; z-index: 1; max-width: 640px; padding-block: 25px; }
.bio-hero__copy .eyebrow { margin-bottom: 24px; }
.bio-hero h1 { margin: 0; font-size: clamp(56px,7vw,86px); font-weight: 550; letter-spacing: -.085em; line-height: .98; }
.bio-hero h1 > span { color: #777774; font-weight: 430; }
.bio-hero__description { max-width: 565px; margin: 24px 0 0; color: #a5a5a1; font-size: 15px; line-height: 1.85; }
.bio-hero__description strong { color: #e4e4e1; font-weight: 590; }
.bio-hero__actions { display: flex; flex-wrap: wrap; gap: 10px; margin-top: 27px; }
.bio-hero__meta { display: flex; flex-wrap: wrap; align-items: center; gap: 9px; margin-top: 32px; color: #797975; font-size: 8px; font-weight: 680; letter-spacing: .11em; }
.bio-hero__meta > i { width: 3px; height: 3px; border-radius: 50%; background: #62625e; }
.bio-hero__meta a { display: inline-flex; align-items: center; gap: 5px; color: #b8b8b4; transition: color .2s ease; }
.bio-hero__meta a:hover { color: #fff; }
.bio-hero__meta .icon { width: 12px; height: 12px; }
.bio-hero__art { position: relative; z-index: 1; width: min(100%,530px); min-height: 366px; justify-self: end; overflow: hidden; border: 1px solid #343431; border-radius: 20px; background: radial-gradient(ellipse at 52% 51%,rgba(255,255,255,.065),transparent 47%),linear-gradient(145deg,#191918,#10100f); box-shadow: 0 32px 75px rgba(0,0,0,.35),inset 0 1px 0 rgba(255,255,255,.055); }
.bio-hero__art::after { position: absolute; right: -70px; bottom: -150px; width: 250px; height: 250px; border: 1px solid rgba(255,255,255,.065); border-radius: 50%; box-shadow: 0 0 0 34px rgba(255,255,255,.015),0 0 0 70px rgba(255,255,255,.01); content: ""; pointer-events: none; }
.bio-hero__art-head,.bio-hero__art-footer { position: absolute; z-index: 2; right: 23px; left: 23px; display: flex; align-items: center; justify-content: space-between; color: #81817d; font-size: 8px; font-weight: 670; letter-spacing: .15em; }
.bio-hero__art-head { top: 20px; padding-bottom: 15px; border-bottom: 1px solid rgba(255,255,255,.09); }
.bio-hero__art-footer { bottom: 20px; justify-content: flex-start; gap: 12px; }
.art-footer-line { width: 52px; height: 1px; background: #555551; }
.bio-hero__art-mark { position: absolute; z-index: 1; right: 23px; bottom: 51px; color: #4b4b47; font-size: 7px; letter-spacing: .14em; }
.energy-diagram { position: absolute; inset: 65px 24px 60px; display: grid; grid-template-columns: 72px minmax(20px,1fr) 82px minmax(20px,1fr) 98px; align-items: center; gap: 6px; }
.energy-diagram__sun,.energy-diagram__leaf,.energy-diagram__atp { position: relative; z-index: 1; display: flex; aspect-ratio: 1; align-items: center; justify-content: center; flex-direction: column; border: 1px solid #454541; border-radius: 50%; background: #161615; box-shadow: 0 8px 28px rgba(0,0,0,.3),inset 0 1px 0 rgba(255,255,255,.07); }
.energy-diagram__sun { color: #e1e1de; }
.energy-diagram__sun-core { font-size: 31px; line-height: 1; }
.energy-diagram__sun small,.energy-diagram__atp small { margin-top: 5px; color: #858581; font-size: 7px; font-weight: 670; letter-spacing: .14em; }
.energy-diagram__leaf { width: 82px; border-radius: 24px; transform: rotate(8deg); }
.energy-diagram__leaf svg { width: 43px; height: 43px; fill: none; stroke: #d5d5d1; stroke-width: 1.45; stroke-linecap: round; stroke-linejoin: round; }
.energy-diagram__atp { width: 98px; aspect-ratio: 1; border-radius: 28px; }
.energy-diagram__atp b { font-size: 25px; font-weight: 620; letter-spacing: -.08em; }
.energy-diagram__connector { position: relative; width: 100%; height: 1px; background: linear-gradient(90deg,transparent,#777773,transparent); }
.energy-diagram__connector::after { position: absolute; top: -3px; right: 0; width: 6px; height: 6px; border-top: 1px solid #aaa; border-right: 1px solid #aaa; content: ""; transform: rotate(45deg); }
.bio-hero__bottom { display: flex; justify-content: space-between; gap: 12px; margin-top: 32px; padding-top: 13px; border-top: 1px solid #272725; color: #686864; font-size: 8px; font-weight: 650; letter-spacing: .14em; }
.topic-hero { padding-top: 36px; }
.topic-hero__art { min-height: 392px; }
.topic-orbit { position: absolute; top: 50%; left: 50%; width: 250px; height: 250px; transform: translate(-50%,-50%); }
.topic-orbit__ring { position: absolute; top: 50%; left: 50%; display: block; border: 1px solid rgba(255,255,255,.12); border-radius: 50%; transform: translate(-50%,-50%); }
.topic-orbit__ring--outer { width: 238px; height: 238px; border-style: dashed; border-color: rgba(255,255,255,.15); }
.topic-orbit__ring--inner { width: 162px; height: 162px; border-color: rgba(255,255,255,.19); }
.topic-orbit__node { position: absolute; z-index: 2; display: grid; width: 47px; height: 47px; place-items: center; border: 1px solid #555551; border-radius: 15px; background: #1a1a19; box-shadow: 0 9px 25px rgba(0,0,0,.35); }
.topic-orbit__node--sun { top: 18px; left: 37px; color: #e5e5e2; font-size: 27px; }
.topic-orbit__node--leaf { right: 12px; bottom: 48px; transform: rotate(8deg); }
.topic-orbit__node--leaf svg { width: 27px; height: 27px; fill: none; stroke: #e0e0dc; stroke-width: 1.45; stroke-linecap: round; stroke-linejoin: round; }
.topic-orbit__core { position: absolute; top: 50%; left: 50%; display: flex; width: 94px; height: 94px; align-items: center; justify-content: center; flex-direction: column; border: 1px solid rgba(255,255,255,.3); border-radius: 28px; background: linear-gradient(145deg,#282827,#111); box-shadow: 0 14px 38px rgba(0,0,0,.4),inset 0 1px 0 rgba(255,255,255,.1); transform: translate(-50%,-50%) rotate(-7deg); }
.topic-orbit__core b { font-size: 24px; font-weight: 700; letter-spacing: -.1em; }
.topic-orbit__core small { margin-top: 2px; color: #969692; font-size: 6px; letter-spacing: .1em; }
.topic-orbit__tag { position: absolute; z-index: 3; padding: 6px 9px; border: 1px solid #3d3d39; border-radius: 7px; background: #151514; color: #a2a29e; font-size: 7px; font-weight: 680; letter-spacing: .12em; }
.topic-orbit__tag--one { top: 45px; right: 17px; }
.topic-orbit__tag--two { bottom: 22px; left: 56px; }

/* Course overview page */
.bio-section-heading { display: flex; align-items: end; justify-content: space-between; gap: 28px; margin-bottom: 35px; }
.bio-section-heading h2,.lesson-content-heading h2 { margin: 0; font-size: clamp(38px,5vw,60px); font-weight: 510; letter-spacing: -.075em; line-height: 1.04; }
.bio-section-heading h2 span,.lesson-content-heading h2 span { color: #777774; font-weight: 430; }
.bio-section-heading .section-description { max-width: 560px; }
.bio-section-heading--split { align-items: end; }
.course-status { display: inline-flex; flex: 0 0 auto; align-items: center; gap: 8px; padding: 9px 12px; border: 1px solid #30302d; border-radius: 999px; color: #93938f; font-size: 8px; font-weight: 680; letter-spacing: .12em; white-space: nowrap; }
.course-status > span { width: 6px; height: 6px; border-radius: 50%; background: #c8c8c4; box-shadow: 0 0 9px rgba(255,255,255,.18); }
.course-topic-section { padding: 91px 0 105px; border-top: 1px solid #1c1c1a; background: #0d0d0c; }
.course-topic-card { display: grid; grid-template-columns: 1.02fr .98fr; overflow: hidden; border: 1px solid #343431; border-radius: 17px; background: linear-gradient(140deg,#181817,#10100f 70%); box-shadow: 0 26px 60px rgba(0,0,0,.22); }
.course-topic-card__main { position: relative; min-height: 440px; padding: clamp(25px,4.5vw,48px); }
.course-topic-card__meta,.course-topic-card__outline-head { display: flex; align-items: center; justify-content: space-between; gap: 10px; color: #81817d; font-size: 8px; font-weight: 680; letter-spacing: .13em; }
.course-topic-card__meta i { color: #555551; font-style: normal; }
.course-topic-card__index { color: #555551; font-size: 10px; }
.course-topic-card__symbol { display: grid; width: 47px; height: 47px; margin-top: 48px; place-items: center; border: 1px solid #3b3b38; border-radius: 13px; background: #1c1c1b; color: #ddd; }
.course-topic-card__symbol svg { width: 27px; height: 27px; fill: none; stroke: currentColor; stroke-width: 1.25; stroke-linecap: round; stroke-linejoin: round; }
.course-topic-card h3 { margin: 19px 0 0; font-size: clamp(36px,5vw,52px); font-weight: 520; letter-spacing: -.07em; line-height: .99; }
.course-topic-card h3 a { transition: color .2s ease; }
.course-topic-card h3 a:hover { color: #c7c7c4; }
.course-topic-card h3 .icon { display: inline-block; width: 20px; height: 20px; margin-left: 5px; color: #a6a6a2; vertical-align: middle; }
.course-topic-card__main > p { max-width: 430px; margin: 16px 0 0; color: #92928e; font-size: 12px; line-height: 1.75; }
.course-topic-card__button { display: inline-flex; align-items: center; gap: 10px; margin-top: 26px; color: #e4e4e1; font-size: 11px; font-weight: 620; }
.course-topic-card__button .icon { width: 15px; height: 15px; }
.course-topic-card__outline { padding: 32px 30px; border-left: 1px solid #2b2b28; background: rgba(0,0,0,.11); }
.course-topic-card__outline-head { padding-bottom: 17px; border-bottom: 1px solid #30302d; }
.course-topic-card__outline ol { display: grid; gap: 2px; margin: 13px 0 0; padding: 0; list-style: none; }
.course-topic-card__outline li a { display: flex; min-height: 38px; align-items: center; gap: 12px; padding: 5px 8px; border-radius: 6px; color: #a5a5a1; font-size: 10px; transition: color .18s ease,background .18s ease; }
.course-topic-card__outline li a:hover { background: rgba(255,255,255,.045); color: #f4f4f2; }
.course-topic-card__outline li a > span { color: #676763; font-size: 8px; font-weight: 650; }
.course-source-note { display: grid; grid-template-columns: 38px minmax(0,1fr) auto; align-items: center; gap: 15px; margin-top: 18px; padding: 16px 18px; border: 1px solid #282826; border-radius: 10px; background: rgba(255,255,255,.014); }
.course-source-note__mark { display: grid; width: 36px; height: 36px; place-items: center; border: 1px solid #393936; border-radius: 9px; color: #d0d0cc; font-size: 8px; font-weight: 730; letter-spacing: .06em; }
.course-source-note p { margin: 0; color: #8f8f8b; font-size: 10px; line-height: 1.6; }
.course-source-note > a,.curriculum-scope-card > a { display: inline-flex; align-items: center; gap: 7px; color: #d0d0cc; font-size: 10px; font-weight: 620; white-space: nowrap; transition: color .2s ease; }
.course-source-note > a:hover,.curriculum-scope-card > a:hover { color: #fff; }
.course-source-note .icon,.curriculum-scope-card > a .icon { width: 14px; height: 14px; }

/* Topic map */
.topic-map-section { padding: 75px 0 89px; border-top: 1px solid #1c1c1a; }
.topic-map-grid { display: grid; grid-template-columns: repeat(2,minmax(0,1fr)); align-items: stretch; gap: 12px; }
.topic-map-card { display: flex; min-width: 0; flex-direction: column; padding: 21px 22px 19px; border: 1px solid #292927; border-radius: 12px; background: linear-gradient(145deg,rgba(255,255,255,.027),rgba(255,255,255,.006)); transition: border-color .2s ease,background .2s ease,transform .2s ease; }
.topic-map-card:hover { transform: translateY(-2px); border-color: #484844; background: linear-gradient(145deg,rgba(255,255,255,.05),rgba(255,255,255,.01)); }
.topic-map-card--last { grid-column: 1 / -1; }
.topic-map-card__top { display: flex; align-items: center; justify-content: space-between; gap: 12px; }
.topic-map-card__number { display: grid; width: 31px; height: 31px; place-items: center; border: 1px solid #3a3a37; border-radius: 8px; background: #181817; color: #d2d2ce; font-size: 9px; font-weight: 660; }
.topic-map-card__type { color: #73736f; font-size: 7px; font-weight: 700; letter-spacing: .14em; }
.topic-map-card h3 { margin: 17px 0 0; color: #e8e8e5; font-size: 17px; font-weight: 570; letter-spacing: -.04em; line-height: 1.35; }
.topic-map-card h3 a { transition: color .18s ease; }
.topic-map-card h3 a:hover { color: #aaa; }
.topic-map-card > p { margin: 7px 0 0; color: #8d8d89; font-size: 10px; line-height: 1.65; }
.topic-map-card__main-link { display: inline-flex; align-items: center; gap: 8px; align-self: flex-start; margin-top: auto; padding-top: 19px; color: #bdbdb9; font-size: 9px; font-weight: 630; }
.topic-map-card__main-link .icon { width: 14px; height: 14px; }
.topic-map-card__links { display: grid; grid-template-columns: repeat(2,minmax(0,1fr)); gap: 3px 14px; margin: 14px 0 0; padding: 13px 0 0; border-top: 1px solid #292927; list-style: none; }
.topic-map-card__links li { min-width: 0; }
.topic-map-card__links a { display: block; padding: 5px 0; color: #8e8e8a; font-size: 9px; line-height: 1.45; transition: color .18s ease; }
.topic-map-card__links a::before { margin-right: 6px; color: #555551; content: "↗"; }
.topic-map-card__links a:hover { color: #f4f4f2; }

/* Long-form topic content */
.lesson-content-section { padding: 86px 0 105px; border-top: 1px solid #1d1d1b; background: #0d0d0c; }
.lesson-content-heading { display: flex; align-items: end; justify-content: space-between; gap: 34px; margin-bottom: 42px; }
.lesson-content-heading h2 { font-size: clamp(36px,4.8vw,55px); }
.lesson-content-heading > p { max-width: 380px; margin: 0 0 3px; color: #8e8e8a; font-size: 11px; line-height: 1.7; }
.lesson-layout { display: grid; grid-template-columns: 210px minmax(0,1fr); align-items: start; gap: clamp(28px,5vw,58px); }
.lesson-sidebar__inner { position: sticky; top: calc(var(--header-height) + 27px); }
.lesson-sidebar__inner > p { display: flex; justify-content: space-between; margin: 0 0 13px; color: #72726e; font-size: 8px; font-weight: 700; letter-spacing: .13em; }
.lesson-sidebar__inner > p span { color: #52524e; }
.lesson-sidebar nav { display: grid; border-top: 1px solid #292927; }
.lesson-sidebar nav a { display: grid; grid-template-columns: 24px minmax(0,1fr); align-items: center; gap: 8px; min-height: 40px; padding: 5px 0; border-bottom: 1px solid #252523; color: #92928e; font-size: 9px; line-height: 1.3; transition: color .18s ease; }
.lesson-sidebar nav a span { color: #62625e; font-size: 8px; }
.lesson-sidebar nav a:hover { color: #f4f4f2; }
.lesson-sidebar__back { display: inline-flex; align-items: center; gap: 7px; margin-top: 17px; color: #858581; font-size: 9px; transition: color .18s ease; }
.lesson-sidebar__back:hover { color: #fff; }
.lesson-sidebar__back .icon { width: 13px; height: 13px; }
.lesson-chapters { min-width: 0; }
.lesson-chapter { padding: 37px 0 42px; border-top: 1px solid #343431; scroll-margin-top: calc(var(--header-height) + 20px); }
.lesson-chapter:first-child { padding-top: 0; border-top: 0; }
.lesson-chapter__heading { display: grid; grid-template-columns: 42px minmax(0,1fr); align-items: start; gap: 14px; margin-bottom: 20px; }
.lesson-chapter__number { display: grid; width: 39px; height: 39px; place-items: center; border: 1px solid #3a3a37; border-radius: 10px; background: #171716; color: #d5d5d1; font-size: 10px; font-weight: 670; }
.lesson-chapter__eyebrow { margin: 2px 0 5px; color: #73736f; font-size: 7px; font-weight: 700; letter-spacing: .14em; }
.lesson-chapter__heading h2 { margin: 0; color: #f0f0ed; font-size: clamp(22px,2.8vw,31px); font-weight: 540; letter-spacing: -.055em; line-height: 1.2; }
.lesson-chapter__grid { display: grid; grid-template-columns: minmax(0,1.18fr) minmax(235px,.82fr); align-items: start; gap: 12px; }
.lesson-chapter__main,.lesson-chapter__aside { display: grid; min-width: 0; gap: 9px; }
.chapter-explanation,.chapter-subtopic,.chapter-note,.pdf-resource { border: 1px solid #292927; border-radius: 10px; background: #121211; }
.chapter-explanation { padding: 18px 19px; }
.field-label { margin: 0 0 9px; color: #777773; font-size: 7px; font-weight: 700; letter-spacing: .14em; }
.chapter-explanation > p:last-child { margin: 0; color: #b5b5b1; font-size: 11px; line-height: 1.78; }
.chapter-subtopics { display: grid; gap: 7px; }
.chapter-subtopic { display: grid; grid-template-columns: 37px minmax(0,1fr); align-items: start; gap: 11px; padding: 13px 15px; scroll-margin-top: calc(var(--header-height) + 20px); }
.chapter-subtopic > span { display: inline-block; padding-top: 3px; color: #696965; font-size: 8px; font-weight: 680; letter-spacing: .05em; }
.chapter-subtopic h3 { margin: 0; color: #dfdfdc; font-size: 11px; font-weight: 630; letter-spacing: -.015em; line-height: 1.5; }
.chapter-subtopic p { margin: 4px 0 0; color: #8e8e8a; font-size: 9px; line-height: 1.7; }
.chapter-visual { margin: 0; overflow: hidden; border: 1px solid #30302d; border-radius: 10px; background: linear-gradient(145deg,#181817,#111110); }
.chapter-visual__canvas { display: grid; min-height: 160px; place-items: center; padding: 14px; background: radial-gradient(ellipse at center,rgba(255,255,255,.045),transparent 70%); color: #c7c7c3; }
.chapter-visual__canvas svg { width: 100%; max-width: 310px; height: auto; overflow: visible; }
.chapter-visual figcaption { display: flex; align-items: center; gap: 8px; padding: 10px 12px; border-top: 1px solid #2a2a27; color: #82827e; font-size: 8px; line-height: 1.45; }
.chapter-visual figcaption span { flex: 0 0 auto; color: #b1b1ad; font-size: 6px; font-weight: 700; letter-spacing: .12em; }
.chapter-visual__canvas--atp { min-height: 160px; align-content: center; gap: 15px; }
.atp-structure { display: flex; align-items: center; justify-content: center; gap: 8px; }
.atp-structure__unit { display: grid; min-width: 55px; min-height: 43px; place-items: center; padding: 5px; border: 1px solid #52524e; border-radius: 9px; background: #1b1b19; color: #e1e1de; font-size: 7px; font-weight: 670; letter-spacing: .08em; }
.atp-structure > i { color: #747470; font-size: 11px; font-style: normal; }
.atp-phosphates { display: flex; align-items: center; gap: 3px; }
.atp-phosphates b { display: grid; width: 26px; height: 26px; place-items: center; border: 1px solid #777773; border-radius: 50%; color: #eee; font-size: 8px; font-weight: 600; }
.atp-structure__caption { color: #71716d; font-size: 7px; font-weight: 680; letter-spacing: .14em; }
.chapter-visual__canvas--equation { gap: 13px; }
.equation-visual { display: flex; flex-wrap: wrap; align-items: center; justify-content: center; gap: 7px; margin: 0; color: #ededeb; font-size: clamp(13px,1.5vw,17px); font-weight: 560; letter-spacing: -.035em; text-align: center; }
.equation-visual > b { color: #777773; font-size: 12px; font-weight: 450; }
.equation-visual__light { padding: 5px 7px; border: 1px solid #464642; border-radius: 6px; color: #bcbcb8; font-size: 8px; letter-spacing: .1em; }
.equation-visual__caption { color: #777773; font-size: 6px; font-weight: 700; letter-spacing: .14em; }
.chapter-note { padding: 14px 15px; border-left: 2px solid #676763; }
.chapter-note .field-label { margin-bottom: 6px; }
.chapter-note p:last-child { margin: 0; color: #a2a29e; font-size: 9px; line-height: 1.7; }
.pdf-resource { display: grid; grid-template-columns: 31px minmax(0,1fr) auto; align-items: center; gap: 9px; padding: 12px; }
.pdf-resource__icon { display: grid; width: 30px; height: 30px; place-items: center; border: 1px solid #3e3e3a; border-radius: 8px; color: #c9c9c5; font-size: 7px; font-weight: 720; letter-spacing: .04em; }
.pdf-resource strong { display: block; color: #d4d4d0; font-size: 9px; font-weight: 630; }
.pdf-resource p { margin: 3px 0 0; color: #787874; font-size: 8px; line-height: 1.5; }
.pdf-resource__status { color: #73736f; font-size: 6px; font-weight: 700; letter-spacing: .1em; white-space: nowrap; }

/* MEB scope callout */
.curriculum-scope-section { padding: 0 0 100px; background: #0d0d0c; }
.curriculum-scope-card { display: grid; grid-template-columns: 45px minmax(0,1fr) auto; align-items: center; gap: 18px; padding: 23px; border: 1px solid #343431; border-radius: 13px; background: linear-gradient(145deg,#171716,#10100f); }
.curriculum-scope-card__mark { display: grid; width: 44px; height: 44px; place-items: center; border: 1px solid #444440; border-radius: 11px; color: #e4e4e1; font-size: 9px; font-weight: 720; letter-spacing: .08em; }
.curriculum-scope-card .field-label { margin-bottom: 5px; }
.curriculum-scope-card h2 { margin: 0; color: #e7e7e4; font-size: 17px; font-weight: 570; letter-spacing: -.035em; }
.curriculum-scope-card p:last-child { max-width: 760px; margin: 6px 0 0; color: #92928e; font-size: 9px; line-height: 1.7; }

@media (max-width: 1020px) {
  .bio-hero__grid { grid-template-columns: minmax(0,1fr) minmax(320px,.9fr); gap: 30px; }
  .bio-hero__art { min-height: 340px; }
  .energy-diagram { inset-inline: 18px; grid-template-columns: 62px minmax(15px,1fr) 72px minmax(15px,1fr) 84px; gap: 4px; }
  .energy-diagram__leaf { width: 72px; }
  .energy-diagram__atp { width: 84px; }
  .lesson-layout { grid-template-columns: 175px minmax(0,1fr); gap: 25px; }
  .lesson-chapter__grid { grid-template-columns: minmax(0,1fr) minmax(215px,.8fr); }
}

@media (max-width: 820px) {
  .bio-hero { padding-top: 31px; }
  .bio-hero__grid { grid-template-columns: minmax(0,1fr); gap: 28px; }
  .bio-hero__copy { max-width: 690px; padding-block: 19px 0; }
  .bio-hero h1 { font-size: clamp(58px,10vw,82px); }
  .bio-hero__art { width: min(100%,600px); min-height: 360px; justify-self: center; }
  .energy-diagram { inset-inline: 32px; grid-template-columns: 72px minmax(20px,1fr) 82px minmax(20px,1fr) 98px; gap: 6px; }
  .energy-diagram__leaf { width: 82px; }
  .energy-diagram__atp { width: 98px; }
  .course-topic-card { grid-template-columns: minmax(0,1fr); }
  .course-topic-card__main { min-height: 390px; }
  .course-topic-card__outline { border-top: 1px solid #2b2b28; border-left: 0; }
  .lesson-content-heading { align-items: flex-start; flex-direction: column; gap: 13px; }
  .lesson-content-heading > p { max-width: 560px; }
  .lesson-layout { grid-template-columns: minmax(0,1fr); gap: 25px; }
  .lesson-sidebar__inner { position: static; }
  .lesson-sidebar__inner > p { margin-bottom: 8px; }
  .lesson-sidebar nav { display: flex; overflow-x: auto; gap: 7px; padding: 9px 0; border-top: 1px solid #292927; border-bottom: 1px solid #292927; scrollbar-width: thin; }
  .lesson-sidebar nav a { display: flex; min-height: 34px; flex: 0 0 auto; gap: 7px; padding: 0 10px; border: 1px solid #2d2d2a; border-radius: 7px; white-space: nowrap; }
  .lesson-sidebar__back { margin-top: 9px; }
  .lesson-chapter__grid { grid-template-columns: minmax(0,1.1fr) minmax(230px,.9fr); }
}

@media (max-width: 600px) {
  .page-breadcrumb { padding-top: 15px; }
  .bio-hero { padding: 25px 0 34px; }
  .bio-hero__grid { gap: 23px; }
  .bio-hero__copy .eyebrow { margin-bottom: 19px; font-size: 8px; }
  .bio-hero h1 { font-size: clamp(50px,13vw,68px); }
  .bio-hero__description { margin-top: 17px; font-size: 12px; line-height: 1.75; }
  .bio-hero__actions { gap: 8px; margin-top: 20px; }
  .bio-hero__meta { gap: 7px; margin-top: 22px; font-size: 7px; letter-spacing: .08em; }
  .bio-hero__meta a { gap: 3px; }
  .bio-hero__art { min-height: 290px; border-radius: 15px; }
  .bio-hero__art-head,.bio-hero__art-footer { right: 15px; left: 15px; font-size: 7px; }
  .bio-hero__art-head { top: 15px; padding-bottom: 11px; }
  .bio-hero__art-footer { bottom: 15px; }
  .bio-hero__art-mark { right: 15px; bottom: 40px; }
  .energy-diagram { inset: 53px 13px 48px; grid-template-columns: 54px minmax(10px,1fr) 62px minmax(10px,1fr) 75px; gap: 3px; }
  .energy-diagram__sun-core { font-size: 23px; }
  .energy-diagram__sun small,.energy-diagram__atp small { font-size: 6px; }
  .energy-diagram__leaf { width: 62px; border-radius: 19px; }
  .energy-diagram__leaf svg { width: 34px; height: 34px; }
  .energy-diagram__atp { width: 75px; border-radius: 22px; }
  .energy-diagram__atp b { font-size: 20px; }
  .energy-diagram__connector::after { width: 5px; height: 5px; }
  .bio-hero__bottom { margin-top: 21px; font-size: 7px; }
  .topic-hero__art { min-height: 300px; }
  .topic-orbit { width: 210px; height: 210px; }
  .topic-orbit__ring--outer { width: 205px; height: 205px; }
  .topic-orbit__ring--inner { width: 140px; height: 140px; }
  .topic-orbit__node--sun { top: 13px; left: 29px; }
  .topic-orbit__node--leaf { right: 3px; bottom: 37px; }
  .topic-orbit__tag--one { top: 37px; right: 8px; }
  .topic-orbit__tag--two { bottom: 15px; left: 44px; }
  .bio-section-heading,.bio-section-heading--split { align-items: flex-start; flex-direction: column; gap: 17px; }
  .bio-section-heading h2 { font-size: clamp(36px,10vw,48px); }
  .bio-section-heading .section-description { font-size: 11px; }
  .course-topic-section { padding: 67px 0 73px; }
  .course-topic-card__main { min-height: 370px; padding: 23px 20px; }
  .course-topic-card__symbol { margin-top: 34px; }
  .course-topic-card h3 { font-size: 43px; }
  .course-topic-card__outline { padding: 24px 19px; }
  .course-topic-card__outline li a { min-height: 36px; font-size: 9px; }
  .course-source-note { grid-template-columns: 35px minmax(0,1fr); gap: 11px; padding: 13px; }
  .course-source-note__mark { width: 33px; height: 33px; }
  .course-source-note > a { grid-column: 2; justify-self: start; }
  .topic-map-section { padding: 61px 0 69px; }
  .topic-map-grid { grid-template-columns: minmax(0,1fr); gap: 9px; }
  .topic-map-card,.topic-map-card--last { grid-column: auto; padding: 17px; }
  .topic-map-card h3 { margin-top: 13px; font-size: 15px; }
  .topic-map-card__links { gap: 2px 10px; margin-top: 11px; padding-top: 9px; }
  .topic-map-card__links a { font-size: 8px; }
  .lesson-content-section { padding: 65px 0 72px; }
  .lesson-content-heading { margin-bottom: 28px; }
  .lesson-content-heading h2 { font-size: 38px; }
  .lesson-content-heading > p { font-size: 10px; }
  .lesson-layout { gap: 17px; }
  .lesson-chapter { padding: 28px 0 32px; }
  .lesson-chapter:first-child { padding-top: 0; }
  .lesson-chapter__heading { grid-template-columns: 35px minmax(0,1fr); gap: 10px; margin-bottom: 14px; }
  .lesson-chapter__number { width: 33px; height: 33px; border-radius: 8px; font-size: 9px; }
  .lesson-chapter__heading h2 { font-size: clamp(20px,6vw,25px); }
  .lesson-chapter__grid { grid-template-columns: minmax(0,1fr); gap: 9px; }
  .chapter-explanation { padding: 15px; }
  .chapter-explanation > p:last-child { font-size: 10px; }
  .chapter-subtopic { grid-template-columns: 31px minmax(0,1fr); gap: 8px; padding: 12px; }
  .chapter-subtopic h3 { font-size: 10px; }
  .chapter-subtopic p { font-size: 8px; }
  .chapter-visual__canvas { min-height: 145px; }
  .chapter-note { padding: 12px; }
  .pdf-resource { grid-template-columns: 29px minmax(0,1fr) auto; padding: 10px; }
  .curriculum-scope-section { padding-bottom: 66px; }
  .curriculum-scope-card { grid-template-columns: 37px minmax(0,1fr); gap: 12px; padding: 16px; }
  .curriculum-scope-card__mark { width: 36px; height: 36px; border-radius: 9px; font-size: 8px; }
  .curriculum-scope-card h2 { font-size: 14px; }
  .curriculum-scope-card > a { grid-column: 2; justify-self: start; }
}

@media (max-width: 360px) {
  .energy-diagram { inset-inline: 9px; grid-template-columns: 49px minmax(8px,1fr) 57px minmax(8px,1fr) 68px; gap: 2px; }
  .energy-diagram__leaf { width: 57px; }
  .energy-diagram__atp { width: 68px; }
  .bio-hero__meta { font-size: 6px; }
  .topic-map-card__links { grid-template-columns: minmax(0,1fr); }
  .course-topic-card__outline li a { font-size: 8px; }
  .atp-structure { gap: 5px; }
  .atp-structure__unit { min-width: 46px; min-height: 38px; font-size: 6px; }
  .atp-phosphates { gap: 2px; }
  .atp-phosphates b { width: 22px; height: 22px; }
  .curriculum-scope-card { grid-template-columns: minmax(0,1fr); }
  .curriculum-scope-card > a { grid-column: auto; }
}